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Rough ray (Raja radula) 🎥 ×16 zoom || Evia Island, Greece 🇬🇷

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The rough ray is named for the spinules that cover its upper surface. In juveniles and males, these rough spinules are only on the snout and the lower half of the tail.
The females of this species have spinules in these areas, but also around the mouth, along the mid-section, and the full tail. These spinules differ from the thorns that are on the inner-eye and from the end of the head to before the dorsal fin. There are two thorns between the dorsal fins.
The subcircular body has a short, curved snout, anterior edges are convex and the extremities are angular wings. Most individuals are around 70 centimeters. The main upper surface colors are light brown and grey, with darker spots and lines. There are two eye-spots with yellow rings and a thick outer darker ring with light dots, placed on either side of the spinal meridian at the level of greatest width of the body. The underside of the ray is white with darkening edges, and some darker spots on the tail.
The females of this species have spinules in these areas, but also around the mouth, along the mid-section, and the full tail. These spinules differ from the thorns that are on the inner-eye and from the end of the head to before the dorsal fin. There are two thorns between the dorsal fins.
The subcircular body has a short, curved snout, anterior edges are convex and the extremities are angular wings. Most individuals are around 70 centimeters. The main upper surface colors are light brown and grey, with darker spots and lines. There are two eye-spots with yellow rings and a thick outer darker ring with light dots, placed on either side of the spinal meridian at the level of greatest width of the body. The underside of the ray is white with darkening edges, and some darker spots on the tail.