Twaite shad (Alosa fallax) migrating upstream to spawn in the River Wye in Wales, May 2020

Taken by Louis Macdonald-Ames, this extraordinary footage shows Twaite shad in the river Wye moving upstream to their spawning grounds in Wales. Otherwise known as "May fish", shad are a member of the herring family that migrate into our rivers to breed in spring. Threatened throughout their range, they are an Annex2 listed species under the EU Habitats Directive.

For more information on this remarkable fish species, please see the Wye & Usk Foundation's website.